I read the whole article. Really interesting. Wish I could grasp the math! Anyway, it seems that ear-tuners, which is what I am, take into account the musicality of Western keyboard music, which is based on equal temperament, so that they instinctively know how to compensate for the errors caused by the underlying waveform conflicts, so that the result sounds "musical." My recollection is that these errors are caused by imperfections in the materials, in this case piano wire, an extremely hard and durable metal (which nevertheless does break. I've broken strings during tuning, and it's not a pleasant experience. I would cringe when stretching those upper notes). Our instructor, Robert Perkins (don't know if he's still in business, I went to his school in the 1980's) definitely had us trust our ears when in doubt. But some ears are better than others. These are the tuners who end up tuning the concert grands for the virtuosi of the performance world. Mastering this process involves internalizing the rates of beat patterns in the various intervals of the temperament octave. for example, one interval might involve beats going at 3 per 5 seconds, another one per five seconds, and so on. How do you memorize this? Well, one trick not involving electronic devises (to which he was vehemently opposed) is to use a metronome. you set it at a certain speed, and memorize that speed. sometimes you have to count every other tick. Then you match the rate the beats are going to that speed you have memorized. Sound difficult? It is. This is why it took me 4 months to finally get it. It's a Zen thing. It has to become part of you. but it's a great feeling when you know you can tune just about any piano with only your ear. (The other thing is tuning it so that it stays in tune a reasonable amount of time. This is called "setting the pins. Another thing that separates the best from the rest).