They're the peak value in dB attained by the associated track. If it reads -6.2, that means the highest level attained by the track, taking the fader setting and any effects into account, was -6.2dB.
You might want to check out my first post in the 31 Days of Tips thread about the meters. There was another post about peak hold, which is what you're seeing.
A reasonable peak value is less than 0 but more than -18. Those are arbitrary numbers, there's no "headroom police."